Scientific commentary
How Giraffes Kept Track as Hidden Food Added Up
A carrot experiment at Barcelona Zoo asked giraffes to follow a meal that changed out of sight—and gave researchers a way to test how they did it.

Explore giraffe behaviour & cognition through original research. Read the questions, methods and findings behind each study, with direct links to its sources.
67 articles
Scientific commentary
A carrot experiment at Barcelona Zoo asked giraffes to follow a meal that changed out of sight—and gave researchers a way to test how they did it.
Scientific commentary
A bird’s warning can cost a giraffe a mouthful of leaves. Playback experiments show that the interruption lasts longer in a landscape shared with lions.
Scientific commentary
Beneath a giraffe’s hoof, several structures share the work of supporting each step. A study after death reveals how damage can involve them together.
Four giraffes chose between hidden samples from food mixtures. Changing the proportions revealed a decision-making ability that simple counting cannot explain.
Observations of 66 giraffes across 18 US zoos linked browsing and extracting food with fewer repetitive mouth behaviours, revealing why feeding time matters.
Tracking giraffes across Africa reveals how plentiful food, patchy resources and human disturbance shape the space animals use and the distances they move.
Giraffes in Uganda used both resident and seasonal movement strategies. Photographs revealed that males were more likely to move between habitats.
Photographic records from Tanzania reveal how movement connects growing and declining giraffe populations, and why counting animals can miss the difference.
A review of fifteen years of giraffe photographs removed duplicate identities and corrected abundance estimates, while still finding strong recovery.
A playback experiment separated sound from sight and found stronger giraffe reactions to vehicle and drone noise at a reserve with less human exposure.
Giraffes at two South African reserves noticed lion recordings quickly, but animals living with lions sustained their vigilance for much longer.
Giraffes outperformed bison and forest buffalos in a small cognition study, but a shaken empty box exposed the importance of checking how they chose.
Observations of 22 giraffe mothers and 47 calves tested seven explanations for shared nursing, finding evidence for reciprocal care and opportunistic calves.
Observers assessed the expressive behaviour of three zoo giraffes and found links between keeper actions and whether animals appeared calm or anxious.
A study followed two giraffes through Oregon Zoo’s 2020 closure and reopening, separating visitor transitions from seasonal and social changes.
Cameras monitored two giraffes during a Christmas light event at Knowsley Safari, finding little change under their existing indoor winter routine.
A small study combined blood measures and body condition to investigate whether accumulated life events leave a measurable physiological signal in giraffes.
GPS records, observations and dung samples revealed how seven giraffes responded to researchers—and why appearing accustomed to people can mislead.
Seven years of photographs showed how young giraffes can leave their original social community while remaining in familiar country in northern Tanzania.
A study of 399 female giraffes linked similar spot shapes to social association, raising a question about recognition without proving how it works.
Networks from two Kenyan populations revealed strong same-sex peer associations in adolescent giraffes, distinguishing repeated company from many contacts.
Five giraffes investigated six scents differently, while a separate herd trial showed why a change in enclosure use needs careful interpretation.
A study of six zoo giraffes compared mapped distances with friendly contact, showing why proximity alone can give an incomplete picture of relationships.
A zoo pilot compared feeding encounters with matched control periods, finding no clear adverse behavioural change while exposing gaps in welfare assessment.
Six years of photographs revealed overlapping giraffe communities in Tanzania and a link between proximity to homesteads and the pattern of female ties.
Three-dimensional scans of giraffe and five other mammal hearts reveal branching muscle groups, changing how researchers picture the pumping wall.
A numerical giraffe model shows how long legs raise the heart and reduce the pumping cost of reaching the same height with an even longer neck.
Walking measurements and skeletal models reveal why a giraffe’s long legs can increase muscular force demands, despite their nearly upright posture.
A more continuous okapi genome revealed striking chromosome differences from giraffes, showing why close relatives need not share the same DNA packaging.
Photographic surveys in Murchison Falls mapped surviving giraffes with snare injuries, revealing uneven exposure without measuring the total death toll.
Tracking six giraffes after translocation distinguished repeated use of a home area from prolonged exploration, offering an early but limited success measure.
A NamibRand study followed thirteen giraffes through sparse habitat, revealing concentrated food use and why a small census cannot establish resilience.
Overnight recordings in three zoos uncovered harmonic giraffe hums, opening a communication question that the sounds alone could not answer.
A field study followed male giraffe sparring, finding size-matched partners and strong individual side preferences without proving future fighting success.
Repeated yielding at feeding time revealed ordered relationships in captive giraffes, with age and stable female ranks complicating an egalitarian image.
Tracking two wild populations showed why simply counting giraffes together can conceal partner preferences that become clearest during foraging.
A two-container experiment showed giraffes locating hidden food after a short wait, while longer delays exposed the limits of this particular memory task.
Giraffes triggered sound by touching or approaching experimental devices, but more activations did not establish that they preferred the sound.
A study across 18 zoos found repetitive behaviours in every observed giraffe, yet their frequency varied enough to change what an average could mean.
Head-mounted sensors distinguished several giraffe behaviours, but testing a new animal exposed the limits of learning from familiar movements.
Sensors followed three wild giraffes through whole nights, revealing two periods of greater rest separated by increased activity around midnight.
Night observations showed some giraffes remaining active while others rested their heads, raising a careful question about protection within a group.
Four kinds of cut branches produced different browsing patterns in five giraffes, and observations after feeding changed how the results were understood.
A summer-to-winter housing comparison found individual behaviour changes without a clear seasonal shift in resting time or hormone measurements.
Laser scans of giraffes, okapis and rhinoceroses show how to compare trunk shape, while exposing the gap between measuring outline and measuring fat.
Electrical recordings and nerve microscopy reveal why long legs leave giraffes with unavoidable delays between a disturbance and a muscle response.
Computer models of Discokeryx tested how reinforced skull and neck joints resisted a head-on collision, separating mechanical evidence from ancestry claims.
Observations at Lake Nakuru found a seasonal shift in one important giraffe food, while the overall share of time spent feeding changed little.
Repeated observations in Hwange showed giraffes eating grass while trees still had leaves, raising a question about nutrients beyond energy.
Serengeti camera traps found oxpeckers sleeping on giraffes and other mammals, revealing a different nighttime pattern from daytime host use.
Observations in two South African parks found zebras scanning less with giraffes nearby and directing more of their attention towards herd mates.
Observations in Namibia connect giraffe urine sampling, a curled lip and the sensory route that may help males find females approaching fertility.
In a Senegalese reserve, giraffe mothers with stronger associations also had calves that spent more time together, revealing a shared social pattern.
Field observations and dung samples in Zimbabwe linked mature male giraffes’ changing courtship activity to hormone changes over days and weeks.
A Knowsley Safari case study followed two male giraffes through an intervention, reduced sparring and a later reversal in their relationship.
Around-the-clock video of three female giraffes showed daytime changes after their group changed, while nighttime routines remained more stable.
A comparison of six laboratory assays showed why measuring hormone traces in giraffe dung needs validation before it can inform welfare decisions.
A video study tested whether focusing on the mouth could help distinguish giraffe licking from standing, walking and eating in a zoo enclosure.
Five years of giraffe sightings in Tanzania distinguished the number and reach of social connections from the strength of close relationships.
A seven-day study at Alipore Zoo found uneven patterns of repetitive licking, showing why a group average can hide individual welfare concerns.
A Kenyan study linked parasite detection to associations outside a giraffe’s usual social circle, revealing why the kind of social connection matters.
Researchers compared giraffe associations with independently measured bacterial types to test what a social network can reveal about microbial sharing.
Two Kenyan populations both showed preferred companions, yet differed in association strength and connectedness under a shared observation method.
Giraffes sometimes chew bones. Field observations and a digestion experiment reveal why the mineral explanation needs a closer look.
Researchers amplified tiny colour changes in a giraffe’s leg to recover a pulse signal from online footage, opening a window on contact-free physiology.
A photographic study in Tanzania tested whether severe skin lesions go with lion-attack marks, revealing how survivors can complicate a predation study.
GPS tracking in Chad found that female Kordofan giraffes used larger wet-season ranges, showing why protection must account for changing access to habitat.